Genetics

Polypeptide Synthesis

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  • Transcription
    • Begins with the binding of transcription factors to DNA within a promoter sequence (such as the TATA box)

    • One strand of DNA forms a template for the synthesis of messenger RNA
    • This is catalysed by RNA polymerase which binds to a DNA promoter region
    • The process moves along the DNA strand as the RNA molecule is built in the 5’ to 3’ direction

    • The process stops when it meets the termination signal on the DNA molecule
    • This transcript is an exact replica of the DNA strand
    • The introns are excised and the exons spliced together at spliceosomes
      • Some molecules undergo ‘alternative’ splicing and thus form other active mRNA 

      • This is another mechanism of regulating gene expression
      • Splicing occurs in the nucleus
  • The mRNA molecule then leaves the nucleus
  • NB: mRNA is monocistronic meaning it only encodes for one protein
  • Its 5’ end is blocked by 7-methylguanine and a poly-adenine tail is added to the 3’ end which aids transport to the cytoplasm and provides stability

  • Translation
    • Performed by specific transfer RNA (tRNA) which is specific for an amino acid and transports them to the mRNA template on the ribosomes

      • Only 1% of cellular RNA is mRNA
    • Amino acids specific for the three nucleotide bases are attached by their C-terminal ends

    • Each tRNA anticodon is linked with the matching codon on the mRNA
    • The process is controlled by aminoacyl-tRNA synthetases which couples amino acids to the matching tRNA molecule and peptidyl transferase which uncouples tRNA from amino acids and joins it to the chain

    • Translation is always initiated at a codon specifying methionine
    • Initiation factor 2 (IF-2) catalyses the reaction between specific tRNA molecules and the mRNA

    • The mRNA is read a codon at a time from the 5’ to 3’ direction
    • The polypeptide chain is built from the N-terminal end to the C-terminal end
    • The stop codon initiates release factor to bind the A site (this is where incoming tRNA molecules bind), blocking further tRNA and releasing the polypeptide into the cytoplasm

    • The polypeptide chain can be further modified after translation

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